Signing pools (delegates) earn a small fraction of the transaction fee from the block. Also, if you have enough stake, you will be proactive about protecting its value.
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I just worry the incentive is not sufficient to maintain sufficient number of nodes for the network.
"if you have enough stake, you will be proactive about protecting its value."
The problem of this statement is how many is "enough". Different people may have different interpretations. Not many of us have a significant stake. Many of us only have a tiny fraction (not enough to move the price if we dump all our shares to any exchange). Do we have the incentive to proactively protect its value and take part in the network? I don't know. (At least, I do not run my bitcoin client or miner programs now)
Unless the security of protocol does not depend on the number of nodes taking part in the network, I guess sufficient incentive is crucial to motivate people to take part. I am not a network expert, I don't know. May be I just don't understand.